ZOOLOGY: STUDY GUIDE - Ye. O. Nevedomska - 2013
LECTURE 10. PHYLUM ANNELIDA, or SEGMENTED WORMS (Annelida)
2. Evolutionary Advancements of the Phylum Annelida
The progressive evolutionary Features of the phylum Annelida include:
1) The Emergence of a secondary body cavity, or coelom, lined with its own epithelial walls;
2) segmentation of the body into metameres containing repeating internal organ structures (metamerism);
3) further advancement of The Nervous system: a significant enlargement of the cerebral ganglion, The formation of a circumoesophageal nerve ring, and a ventral nerve cord resulting from the fusion of lateral Nerve Trunks and The Development of ganglia in each segment;
4) the appearance of a Circulatory system, as well as dermal gills in the Respiratory system, which substantially increased The rate of METABOLISM;
5) greater complexity of the Digestive System (Differentiation of the midgut into specialized regions, allowing for continuous Digestion);
6) the development of paired body outgrowths on each trunk segment — parapodia (from the Greek para — beside, and podion — FOOT), which function as locomotory Organs;
7) the formation of a multicellular excretory system — metanephridia, occurring as a pair in each body segment.
Last update: 14/08/2026
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